国家传染病防治科技重大专项(2008ZX10002-011、2012ZX10004-503);科技部国际科技合作专项(2011DFA31030);国家自然科学基金(81201289)@@@@sNational major special projects for the prevention and treatment of infectious diseases (2008ZX10002-011、2012ZX10004-503); International Science and technology cooperation project of Ministry of science and technology (2011DFA31030);The National Natural Science Foundation of China (81201289)